By Alexander Marrow

MOSCOW (Reuters) -Russia’s communications regulator on Wednesday demanded an explanation from Apple after applications operated by the Russian state-controlled tech firm VK were removed from the U.S. firm’s App Store.

The regulator, Roskomnadzor, has been a thorn in the side of foreign tech firms, issuing fines over data storage violations and for failures to remove content Russia deems illegal – disputes that have escalated since Moscow sent its armed forces into Ukraine in February.

VK runs Vkontakte, which with over 75 million monthly users is Russia’s largest social network, often compared to Facebook. Along with VK’s email service, Vkontakte must be pre-installed on mobile devices sold in Russia.

Roskomnadzor said Apple’s actions had deprived millions of Russians of access to VK apps and demanded that it explain its decision.

Apple said it follows laws in the jurisdictions where the company operates. Apple said the apps in question were being distributed by developers majority-owned or majority-controlled by one or more parties sanctioned by the UK government.

Regardless of location, the apps cannot be downloaded from any App store, said Apple, which terminated the developer accounts associated with the apps to comply with UK sanctions.

State-controlled VK tightened its grip on Russia’s internet space this month by finalising a deal to buy rival Yandex’s news aggregator, content platform Zen and homepage.

VK CEO Vladimir Kiriyenko is under UK sanctions, along with his father Sergei Kiriyenko, who serves as President Vladimir Putin’s first deputy chief of staff.

Roskomnadzor called the restrictions “discriminatory” and said they violated Russian internet users’ right to free information and communication.

Russia has, however, itself limited its citizens’ access to information by forcing all significant independent Russian media to close and blocking U.S. social media platforms Facebook, Instagram and Twitter.

VK on Tuesday said that its apps, including Vkontakte, continued to work if already installed on smartphones, and that it was continuing to develop and support apps for Apple’s iOS operating system.

Roskomnadzor also asked Apple to explain its removal of other apps, including those of airline Aeroflot and a number of banks such as top lenders Sberbank and VTB.

(Reporting by Alexander Marrow; Editing by Kevin Liffey and Mark Trevelyan)